Calculus for the Life Sciences: A Modeling Approach

Volume I. The First Year Course 

Volume II. Difference and Differential Equations

James L. Cornette and Ralph A. Ackerman, Departments of Mathematics and
Ecology, Evolution, and Organismal Biology,
Iowa State University, Ames, Iowa 50011

You are welcome to use this text at no cost on a non-commercial basis. You may use the text on this web site or download to your computer the text or those chapters of interest.   You may print copies for yourself.  Instructors may print copies for their students and charge the students the printing costs. 

The text is issued under a license by Creative Commons, however, that does not allow you to alter, transform or build upon this work. Please read License

Previous issuance under a license, Creative Commons: by-nc-sa, that allowed one to alter, transform, or build upon the work is now withdrawn, August 4, 2013.

Volume I is appropriate for students who have prepared for but have not studied calculus.

Volume II is suitable for students who have studied calculus and are interested in difference and differential equation models in the life sciences.


About Calculus for the Life Sciences




Descriptions of Volumes I and II


The Authors


Volume I by Chapters


Volume II by Chapters


Volume I, Complete


Volume II Complete






Suggested Solutions to
Volume I Exercises


Suggested Solutions appear at the end of each chapter of both volumes on the web.

This text is reviewed in the MAA Mathematical Sciences Digital Library.

Duane Nykamp of the University of Minnesota has a website, Math Insight, which is a collection of webpages and applets designed to shed light on concepts underlying a few topics in mathematics. Some of his pages (see links) are based on our material.

I would be very pleased to have your comments about this work.

James L. Cornette,  515-292-4344